Weber R. (2008) Phlebopteris (Matoniaceae) en el Triasico y Jurasico de Mexico // in Weber R. (ed.), Plantas triasicas y jurasicas de Mexico: Universidad Nacional Autonoma de Mexico, Instituto de Geologia, Boletin 115, Parte 2, p. 85-115. pdf:http://www.geologia.unam.mx/publicaciones/b115/Bol_115_2.pdf
The oldest known Laurasian Matoniaceae, genus Phlebopteris, are Mid/Late Triassic (late Ladinian/early Carnian) in age. The Carnian (and, perhaps, Norian?) Santa Clara Formation, Central Sonora, Mexico, bears three species of Phlebopteris. Ph. smithii is moderately common. Phlebopteris crenulata sp. nov. and Phlebopteris angustiloba were found together in one Sonoran site. The geographically and stratigraphically close Chinle Formation, SW USA, bears two species, Phlebopteris smithii and Phlebopteris utensis. The Chinle and Santa Clara Formations together are the oldest known center of diversity of the genus. In addition, Phlebopteris otongensis sp. nov. is described from the Early Jurassic (Hettangian) Huayacocotla Formation near Otongo, Hidalgo State, eastern Mexico.